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Gonocephalus
bornensis (Schlegel, 1848):
Lophyurus Bornensis
H. Schlegel. 1848. Bijdr. Dierk. 1: 6; Pl. III; Fig. 2.
Tiaris miotympanum
A. C. L. Günther. 1872. Proc. Zool. Soc. London 1872: 592; Pl. XXXVII; Fig. B.
(type locality: “Labuan” (= Pulau Labuan, 05° 19’N; 115° 13’E; a
Federal Territory off eastern Sarawak, East Malaysia; Borneo; BMNH 1946.8.19.82
and 1946.8.14.64 (two syntypes).
Gonyocephalus borneensis G.
A. Boulenger. 1885. Cat. Lizards British Mus. 1: 288.
Gonyocephalus miotympanum E.
Bartlett. 1895. J. Str. Br. Royal Asiatic Soc. (28): 84.
Gonyocephalus borneensis E.
Bartlett. 1895. J. Str. Br. Royal Asiatic Soc. (28): 84.
Gonyocephalus borneensis T.
Barbour. 1912. Mem. Mus. Comp. Zool. 44(1): 181.
Gonyocephalus miotympanum T.
Barbour. 1912. Mem. Mus. Comp. Zool. 44(1): 181.
Gonyocephalus borneensis N.
De Rooij. 1915. Reptiles Indo-Australian Arch. 1: 106.
Gonyocephalus miotympanum N.
De Rooij. 1915. Reptiles Indo-Australian Arch. 1: 106.
Gonocephalus bellii H.
Wermuth. 1967. Das Tierreich 86: 57 (in part)
Gonocephalus miotympanum H.
Wermuth. 1967. Das Tierreich 86: 60.
Gonocephalus miotympanum K.
R. G. Welch, P. S. Cooke & A. S. Wright. 1990. Lizards Orient: 45.
Gonocephalus denzeri
U. Manthey. 1991. Salamandra 27(3): 152; Figs. 1-3. (type locality: “Kapit. 3.
Division, Sarawak, Borneo (Ost-Malaysia)” (02° 00’N; 112° 54’E); ZMK
50527 [formerly CMNH 188626; holotype.]; ZMK 50528 (formerly CMNH 188623;
paratype). Synonymy by Manthey and Grossmann (1997: 505).
Gonyocephalus borneensis U.
Manthey & W. Grossmann. 1992. Sauria, Berlin 14(3): 12.
Gonocephalus bellii F.
L. Tan. 1993. Checklist Lizards Borneo: 4. (in part)
Gonocephalus borneensis U.
Manthey & W. Grossmann. 1997. Amphibien & Reptilien Südostasiens: 191.
Type
locality: RMNH 3043-44 (two syntypes; fide Manthey and Denzer, 1992e), from
“l’ile de Bornéo”.
Distribution:
Endemic to Borneo (Manthey and Grossmann, 1997: 180).
Localities
in Borneo: SARAWAK: Mount Poi (Smith, 1925b); Kapit, Third Division (Manthey,
1991); Gunung Mulu National Park (Ota et al., 1992: as Gonocephalus miotympanum); Baram River (BMNH 1921.4.1.102; ZRC
2.915); Balingean (BMNH 1904.7.19.45); Kuching (BMNH 1911.1.30.14); Kidi
District, Belaga (BMNH 1933.20.18); Mount Dulit (BMNH 1900.1.23.11-12); LABUAN:
Labuan (BMNH 1946.8.19.82 and 1946.8.14.64); BRUNEI: Batu Apoi (Das, 1995, based
on UBD 333, 356, 480, 507, 515, 576, 608, 686); SABAH: Mount Kinabalu (Manthey
and Denzer, 1982a), as well as more specific localities such as Kiau, Mount
Kinabalu (ZRC 2.888-89); Kaingeran, Mount Kinabalu (BMNH 1960.1.4.15-18);
Merabah (BMNH 1913.3.6.33-36); Sawah (BMNH 1898.7.20.1); Poring Hot Springs
(Malkmus, 1992: as Gonocephalus
miotympanum; Malkmus, 1996; Manthey and Grossmann, 1997: 180); Brumas, 75 km
NW Tawau (ANSP 28653-55); Sandakan Bay (Lidth de Jeude, 1893: as Gonocephalus
miotympanum); Samarang River (BMNH 1974.3832); Kampung Bharu Jumpa (BMNH
1974.3833); KALIMANTAN: Sampit (ZMA 9 067); Samarinda (ZMA 11 585); unlocated:
Gunung Alang (ZRC 2.930).
Derivation:
Latin implying an inhabitant of Borneo.
Remarks:
Gonocephalus borneensis (Schlegel,
1848) was synonymised under Gonocephalus
bellii (Duméril and Bibron, 1837) by Smith (1935: 134), but revived as a
distinct species by Manthey and Grossmann (1992). The taxon has frequently been
misspelt as borneensis (e.g.,
Boulenger, 1885: 288; De Rooij, 1915: 106; Tanner, 1953; Manthey and Grossmann,
1997: 191).
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